The integrated stress response promotes immune evasion through lipocalin 2

Интегрированный стрессовый ответ способствует уклонению от иммунного надзора посредством липокалина 2
Aristotelis Tsirigos, Francisco J. Sánchez‐Rivera, Iannis Aifantis, Thales Papagiannakopoulos, Jozef P. Bossowski, Ray Pillai, John Kilian, Angela Wong Lau, Mari Nakamura, Ali Rashidfarrokhi, Yuan Hao, Ruxuan Li, Katherine Wu, Takamitsu Hattori, Eliezra Glasser, Akiko Koide, Lidong Wang, André M. Leite-Moreira, Cristina Hajdu, Sahith Rajalingam, Sarah E. LeBoeuf, Hortense Le, So Yeon Lee, Jin Woo Oh, Cheolyong Joe, Hyemin Kim, Chan-Young Ock, Se-Hoon Lee, Hao Wang, Angana A. H. Patel, Volkan I. Sayin, Kwok-Kin Wong, Sergei B. Koralov, Mario Pende, Diane M. Simeone, Ioannis K. Zervantonakis, Shohei Koide, Akiko Koide, Shohei Koide, Volkan I. Sayin, Jozef P. Bossowski, Ray Pillai, Diane M. Simeone, Mario Pende, Aristotelis Tsirigos, Seungeun Lee, Angana A.H. Patel, Seungeun Lee
2026-02-18

ATF4–LCN2 axisT cell exclusionimmune evasionintegrated stress responselipocalin 2
Cancer cells activate the integrated stress response (ISR) to adapt to stress and resist therapy1. ISR signals converge on activating transcription factor 4 (ATF4), which controls cell-intrinsic transcriptional programs that are involved in metabolic adaptation, survival and growth2,3. However, whether the ISR–ATF4 axis influences anti-tumour immune responses remains mostly unknown. Here we show that loss of ATF4 decreases tumour progression considerably in immunocompetent mice, but not in immunocompromised ones, by enhancing T cell-dependent anti-cancer immune responses. An unbiased genetic screen of ATF4-regulated genes identifies lipocalin 2 (LCN2) as the principal ATF4-dependent effector that impairs anti-tumour immunity by favouring infiltration with immunosuppressive interstitial macrophages. Furthermore, we find that LCN2 promotes T cell exclusion and immune evasion in preclinical mouse models, and correlates with decreased T cell infiltration in patients with lung and pancreatic adenocarcinomas. Anti-LCN2 antibodies promote robust anti-tumour T cell responses in mouse models of aggressive solid tumours. Our study shows that the ATF4–LCN2 axis has a cell-extrinsic role in suppressing anti-cancer immunity, and could pave the way for an immunotherapy approach that targets LCN2. The transcription factor ATF4 and its effector lipocalin 2 (LCN2) have a key role in immune evasion and tumour progression, and targeting the ATF4–LCN2 axis might provide a way to treat several types of solid tumour by increasing anti-cancer immunity.
1
A genetic screen identifies lipocalin 2 (LCN2) as the principal ATF4-dependent effector suppressing anti-tumour immune responses.
2
ATF4-driven LCN2 promotes immunosuppressive interstitial macrophage infiltration, T cell exclusion and immune evasion in preclinical tumour models.
3
Anti-LCN2 antibodies induce robust anti-tumour T cell responses in mouse models, supporting the ATF4–LCN2 axis as an immunotherapy target.
4
LCN2 expression correlates with decreased T cell infiltration in human lung and pancreatic adenocarcinomas.
5
Loss of ATF4 substantially reduces tumour progression in immunocompetent, but not immunocompromised, mice by enhancing T cell-dependent anti-tumour immunity.

Tumours and tumour–immune interactions governed by the ATF4–LCN2 axis

The ATF4–LCN2-mediated mechanisms of immune evasion, including immunosuppressive macrophage infiltration and T cell exclusion, and their effects on tumour progression and anti-tumour immunity

Publication Details
Publication Date
2026-02-18
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Authors
Aristotelis Tsirigos
Francisco J. Sánchez‐Rivera
Iannis Aifantis
Thales Papagiannakopoulos
Jozef P. Bossowski
Ray Pillai
John Kilian
Angela Wong Lau
Mari Nakamura
Ali Rashidfarrokhi
Yuan Hao
Ruxuan Li
Katherine Wu
Takamitsu Hattori
Eliezra Glasser
Akiko Koide
Lidong Wang
André M. Leite-Moreira
Cristina Hajdu
Sahith Rajalingam
Sarah E. LeBoeuf
Hortense Le
So Yeon Lee
Jin Woo Oh
Cheolyong Joe
Hyemin Kim
Chan-Young Ock
Se-Hoon Lee
Hao Wang
Angana A. H. Patel
Volkan I. Sayin
Kwok-Kin Wong
Sergei B. Koralov
Mario Pende
Diane M. Simeone
Ioannis K. Zervantonakis
Shohei Koide
Akiko Koide
Shohei Koide
Volkan I. Sayin
Jozef P. Bossowski
Ray Pillai
Diane M. Simeone
Mario Pende
Aristotelis Tsirigos
Seungeun Lee
Angana A.H. Patel
Seungeun Lee
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