Nuclear ribosomal internal transcribed spacer (ITS) region as a universal DNA barcode marker for Fungi

Внутренний транскрибируемый спейсерный (ITS) регион ядерной рибосомной ДНК как универсальный маркер ДНК-штрихкодирования грибов
Conrad L. Schoch, Keith A. Seifert, Sabine M. Huhndorf, Vincent Robert, John L. Spouge, C. André Lévesque, Wen Chen, Elena Bolchacova, Kerstin Voigt, P.W. Crous, Andrew N. Miller, Michael J. Wingfield, M. Catherine Aime, Kwang-Deuk An, Feng‐Yan Bai, Robert W. Barreto, Dominik Begerow, Marie‐Josée Bergeron, Meredith Blackwell, Teun Boekhout, Mesfin Bogale, Nattawut Boonyuen, Ana Rosa Burgaz, Bart Buyck, Lei Cai, Qing Cai, Gianluigi Cardinali, Priscila Chaverrí, B. J. Coppins, Ana Crespo, Pilar Cubas, Craig Cummings, Ulrike Damm, Z. Wilhelm de Beer, Sybren de Hoog, Ruth Del‐Prado, Bryn T. M. Dentinger, Javier Diéguez‐Uribeondo, Pradeep K. Divakar, Brian Douglas, Margarita Dueñas, Tuan A. Duong, Ursula Eberhardt, Joan E. Edwards, Mostafa S. Elshahed, K. Fliegerová, Manohar R. Furtado, Miguel A. Garcı́a, Zai-Wei Ge, Gareth Griffith, Kate Griffiths, J.Z. Groenewald, Marizeth Groenewald, Martín Grube, Marieka Gryzenhout, Liang‐Dong Guo, Ferry Hagen, Sarah Hambleton, Richard C. Hamelin, Karen Hansen, Paul Harrold, Gregory Heller, Cesar S. Herrera, Kazuyuki Hirayama, Yuuri Hirooka, Hsiao-Man Ho, Kerstin Hoffmann, Valérie Hofstetter, Filip Högnabba, Peter M. Hollingsworth, Seung‐Beom Hong, Kentaro Hosaka, Jos Houbraken, Karen W. Hughes, Seppo Huhtinen, Kevin D. Hyde, Timothy Y. James, Eric M. Johnson, Joan E. Johnson, Peter R. Johnston, E. B. Gareth Jones, Laura J. Kelly, Paul M. Kirk, Dániel G. Knapp, Urmas Kõljalg, Gábor M. Kovács, Cletus P. Kurtzman, Sara Landvik, Steven D. Leavitt, Audra S. Liggenstoffer, Kare Liimatainen, Lorenzo Lombard, Janet Jennifer Luangsa-ard, H. Thorsten Lumbsch, Harinad B. Maganti, Sajeewa S. N. Maharachchikumbura, María P. Martín, Tom W. May, Alistair R. McTaggart, Andrew S. Methven
2012-03-27

barcode gapfungal DNA barcodinginternal transcribed spacer (ITS)nuclear ribosomal RNA cistronspecies identification
Six DNA regions were evaluated as potential DNA barcodes for Fungi, the second largest kingdom of eukaryotic life, by a multinational, multilaboratory consortium. The region of the mitochondrial cytochrome c oxidase subunit 1 used as the animal barcode was excluded as a potential marker, because it is difficult to amplify in fungi, often includes large introns, and can be insufficiently variable. Three subunits from the nuclear ribosomal RNA cistron were compared together with regions of three representative protein-coding genes (largest subunit of RNA polymerase II, second largest subunit of RNA polymerase II, and minichromosome maintenance protein). Although the protein-coding gene regions often had a higher percent of correct identification compared with ribosomal markers, low PCR amplification and sequencing success eliminated them as candidates for a universal fungal barcode. Among the regions of the ribosomal cistron, the internal transcribed spacer (ITS) region has the highest probability of successful identification for the broadest range of fungi, with the most clearly defined barcode gap between inter- and intraspecific variation. The nuclear ribosomal large subunit, a popular phylogenetic marker in certain groups, had superior species resolution in some taxonomic groups, such as the early diverging lineages and the ascomycete yeasts, but was otherwise slightly inferior to the ITS. The nuclear ribosomal small subunit has poor species-level resolution in fungi. ITS will be formally proposed for adoption as the primary fungal barcode marker to the Consortium for the Barcode of Life, with the possibility that supplementary barcodes may be developed for particular narrowly circumscribed taxonomic groups.
1
A multinational, multilaboratory consortium evaluated six DNA regions as potential universal barcode markers across the fungal kingdom.
2
Compared with ITS, the large subunit provided superior resolution in some groups, while the small subunit showed poor species-level resolution; ITS was recommended as the primary fungal barcode.
3
Mitochondrial cytochrome c oxidase subunit 1 was excluded because fungal amplification is difficult, introns are common, and variability can be insufficient.
4
Protein-coding markers sometimes achieved higher identification accuracy than ribosomal regions, but poor PCR amplification and sequencing success prevented universal use.
5
Supplementary barcodes may be needed for narrowly circumscribed fungal taxonomic groups.
6
The nuclear ribosomal ITS region showed the highest overall identification success and the clearest barcode gap between interspecific and intraspecific variation.

Fungi and candidate DNA barcode regions, particularly the nuclear ribosomal internal transcribed spacer (ITS) region

The comparative identification performance, amplification and sequencing success, species resolution, and barcode-gap characteristics of candidate regions for universal fungal DNA barcoding

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Publication Date
2012-03-27
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Authors
Conrad L. Schoch
Keith A. Seifert
Sabine M. Huhndorf
Vincent Robert
John L. Spouge
C. André Lévesque
Wen Chen
Elena Bolchacova
Kerstin Voigt
P.W. Crous
Andrew N. Miller
Michael J. Wingfield
M. Catherine Aime
Kwang-Deuk An
Feng‐Yan Bai
Robert W. Barreto
Dominik Begerow
Marie‐Josée Bergeron
Meredith Blackwell
Teun Boekhout
Mesfin Bogale
Nattawut Boonyuen
Ana Rosa Burgaz
Bart Buyck
Lei Cai
Qing Cai
Gianluigi Cardinali
Priscila Chaverrí
B. J. Coppins
Ana Crespo
Pilar Cubas
Craig Cummings
Ulrike Damm
Z. Wilhelm de Beer
Sybren de Hoog
Ruth Del‐Prado
Bryn T. M. Dentinger
Javier Diéguez‐Uribeondo
Pradeep K. Divakar
Brian Douglas
Margarita Dueñas
Tuan A. Duong
Ursula Eberhardt
Joan E. Edwards
Mostafa S. Elshahed
K. Fliegerová
Manohar R. Furtado
Miguel A. Garcı́a
Zai-Wei Ge
Gareth Griffith
Kate Griffiths
J.Z. Groenewald
Marizeth Groenewald
Martín Grube
Marieka Gryzenhout
Liang‐Dong Guo
Ferry Hagen
Sarah Hambleton
Richard C. Hamelin
Karen Hansen
Paul Harrold
Gregory Heller
Cesar S. Herrera
Kazuyuki Hirayama
Yuuri Hirooka
Hsiao-Man Ho
Kerstin Hoffmann
Valérie Hofstetter
Filip Högnabba
Peter M. Hollingsworth
Seung‐Beom Hong
Kentaro Hosaka
Jos Houbraken
Karen W. Hughes
Seppo Huhtinen
Kevin D. Hyde
Timothy Y. James
Eric M. Johnson
Joan E. Johnson
Peter R. Johnston
E. B. Gareth Jones
Laura J. Kelly
Paul M. Kirk
Dániel G. Knapp
Urmas Kõljalg
Gábor M. Kovács
Cletus P. Kurtzman
Sara Landvik
Steven D. Leavitt
Audra S. Liggenstoffer
Kare Liimatainen
Lorenzo Lombard
Janet Jennifer Luangsa-ard
H. Thorsten Lumbsch
Harinad B. Maganti
Sajeewa S. N. Maharachchikumbura
María P. Martín
Tom W. May
Alistair R. McTaggart
Andrew S. Methven
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