Immunizations with diverse sarbecovirus receptor-binding domains elicit SARS-CoV-2 neutralizing antibodies against a conserved site of vulnerability

Colin J. Jackson, Philip M. Hansbro, Hans‐Martin Jäck, Edith Roth, Anthony D. Kelleher, Simon T. Schafer, Robert Brink, Anupriya Aggarwal, Stuart Turville, William D. Rawlinson, Matt D. Johansen, Nicole G. Hansbro, Romain Rouet, Thomas Winkler, Christopher C. Goodnow, Hui Li, Helen Lenthall, Matthew A. Spence, Alastair G. Stewart, Sebastian Schulz, Warwick J. Britton, Katherine Jackson, Rowena A. Bull, Mandeep Singh, Peter Schofield, Daniel Christ, Sean Emery, Jake Y. Henry, Gregory J. Walker, Harikrishnan Balachandran, Deborah L. Burnett, David B. Langley, Alberto Ospina Stella, Bernadette M. Saunders, Jennifer Jackson, Franka Witthauer, Claire Milthorpe, Marianne Martinello, Rudolfo Karl
2021-10-29

SCID:  54.1/yyabew43
Viral mutations are an emerging concern in reducing SARS-CoV-2 vaccination efficacy. Second-generation vaccines will need to elicit neutralizing antibodies against sites that are evolutionarily conserved across the sarbecovirus subgenus. Here, we immunized mice containing a human antibody repertoire with diverse sarbecovirus receptor-binding domains (RBDs) to identify antibodies targeting conserved sites of vulnerability. Antibodies with broad reactivity against diverse clade B RBDs targeting the conserved class 4 epitope, with recurring IGHV/IGKV pairs, were readily elicited but were non-neutralizing. However, rare class 4 antibodies binding this conserved RBD supersite showed potent neutralization of SARS-CoV-2 and all variants of concern. Structural analysis revealed that the neutralizing ability of cross-reactive antibodies was reserved only for those with an elongated CDRH3 that extends the antiparallel beta-sheet RBD core and orients the antibody light chain to obstruct ACE2-RBD interactions. These results identify a structurally defined pathway for vaccine strategies eliciting escape-resistant SARS-CoV-2 neutralizing antibodies.
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2021-10-29
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Authors
Colin J. Jackson
Philip M. Hansbro
Hans‐Martin Jäck
Edith Roth
Anthony D. Kelleher
Simon T. Schafer
Robert Brink
Anupriya Aggarwal
Stuart Turville
William D. Rawlinson
Matt D. Johansen
Nicole G. Hansbro
Romain Rouet
Thomas Winkler
Christopher C. Goodnow
Hui Li
Helen Lenthall
Matthew A. Spence
Alastair G. Stewart
Sebastian Schulz
Warwick J. Britton
Katherine Jackson
Rowena A. Bull
Mandeep Singh
Peter Schofield
Daniel Christ
Sean Emery
Jake Y. Henry
Gregory J. Walker
Harikrishnan Balachandran
Deborah L. Burnett
David B. Langley
Alberto Ospina Stella
Bernadette M. Saunders
Jennifer Jackson
Franka Witthauer
Claire Milthorpe
Marianne Martinello
Rudolfo Karl
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